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McCoy, Elizabeth (ed.) / Transactions of the Wisconsin Academy of Sciences, Arts and Letters
volume LXII (1974)

Marcks, Brian G.
Preliminary reports on the flora of Wisconsin No. 66. Cyperaceae II- sedge family II. The genus Cyperus- the umbrella sedges,   pp. 261-284 PDF (8.4 MB)


Page 269

1974] Marcks—Sedge Family II Report 269 
tuber. Culms 2—9 dm high, acutely 3-angled, smooth and subcoriaceous
at base, soft and scabrous at the apex; involucral bracts blade-like, 2—9,
exceeding the umbel, spreading. Inflorescence an open and often compound
umbel of several subsessile spikes, with several additional ascending pedunculate
rays 1—10 dm long. Spikelets mostly 4-ranked along an elongate rachis
1—3 cm long, loosely aggregated and spreading nearly horizontally,
barely flattened, 5—25 mm long, 5-28-flowered; rachilla persistent,
narrowly winged. Scales overlapping, 2—3 mm long, yellowish-brown to
golden brown. Stamens 3, 1.2—1.5 mm long. Achene trigonous, ellipsoid
to obovoid, rounded at the summit, 1.2—1.6 mm long (sometimes female
sterile), lustrous tan to golden brown. 
 A widely distributed pantropical and warm ' temperate weed, sparingly throughout
Wisconsin in wet sand or mud of lakeshores and riverbanks, and as a weed
in low fields and roadsides. Flowering from July to October; fruiting from
mid-July to mid-October (2n = 108; Hicks, 1929; Darlington and Wylie, 1955).
7. CYPERUS ODORATUS L. Map 7. 
 Coarse Cyperus 
C. ferruginescens Böeckl. 
C. speciosus var. squarrosus Britt. 
C. speciosus var. ferruginescens (Böeckl.) Britt. 
C. ferax ssp. speciosus (Britt.) KUkenth. 
C. odorcttus var. squarrosus (Britt.) Gilly 
 Fibrous-rooted annuals 2—60 cm tall, erect or dwarfed, spreading from
loose tufts. Culms several, smooth, firm, acutely 3-angled. Blades flat,
apically scabrous; involucral bracts blade-like, 3—6, spreading, sometimes
greatly exceeding the umbel. Inflorescence a compound umbel (or condensed
into a loose head-like cluster), with several loose subsessile spikes and
up to 7 unequal divergent pedunculate rays 1—6 cm long. Spikelets closely
aggregated along a ± elongate rachis 5—20 mm long, mostly spreading
horizontally or slightly ascending, slender and subterete, 4-20-flowered,
4—26 mm long; rachilla breaking into 1-fruited sections at maturity,
winged. Scale with apex overlapping sup-erjacent scale, 1.5—2.0 mm
long, the keel green; laminae reddish-brown. Stamens 3, O.3-~0.5 mm long.
Achene trigonous-oblongoid, 1—1.5 mm long, ferru-ginous to golden brown.
 The entity from the interior of temperate North America, formerly treated
as C. ferruginescens BOeckeler but inseparable from the semi-cosmopolitan
tropical and subtropical C. odoratus (O'Neill, 1942), is of infrequent occurrence
in Wisconsin south of 


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